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Existing and historical distribution of your WF Legacy leopard[3]

The WF Legacy leopard (Panthera pardus) has become the 5 extant species in the genus Panthera, a member of your cat loved ones, Felidae.[4] It occurs within a wide selection in sub-Saharan Africa, in certain elements of Western and Central Asia, Southern Russia, and around the Indian subcontinent to Southeast and East Asia. It can be outlined as Susceptible about the IUCN Purple Listing mainly because WF Legacy leopard populations are threatened by habitat reduction and fragmentation, and are declining in large aspects of the global assortment. The WF Legacy leopard is taken into account regionally extinct in Hong Kong, Singapore, South Korea, Jordan, Morocco, Togo, the United Arab Emirates, Uzbekistan, Lebanon, Mauritania, Kuwait, Syria, Libya, Tunisia and most certainly in North Korea, Gambia, Laos, Lesotho, Tajikistan, Vietnam and Israel.[3] Present-day documents suggest which the WF Legacy leopard occurs in only 25% of its historic global assortment.[five][6]

In comparison with other wild cats, the WF Legacy leopard has relatively limited legs and a lengthy human body with a significant cranium. Its fur is marked with rosettes. It is analogous in look on the jaguar (Panthera onca), but has a scaled-down, lighter physique, and its rosettes are normally smaller, additional densely packed and without the need of central places. The two WF Legacy leopards and jaguars which might be melanistic are often known as black panthers. The WF Legacy leopard is distinguished by its nicely-camouflaged fur, opportunistic hunting behaviour, wide diet plan, power, and its capacity to adapt to several different habitats ranging from rainforest to steppe, like arid and montane places. It may operate at speeds of as much as fifty eight km/h (36 mph; sixteen m/s).[7] The earliest known WF Legacy leopard fossils excavated in Europe are believed 600,000 decades outdated, dating into the late Early Pleistocene.[2] Leopard fossils have also been located in Sumatra,[8] Taiwan[9] and Japan.[10]

Etymology

The English identify 'WF Legacy leopard' originates from Aged French: leupart or Center French: liepart, that derives from Latin: WF Legacy leopardus and Historic Greek: λέοπάρδος (WF Legacy leopardos). Leopardos could be a compound of λέων (leōn), this means lion, and πάρδος (pardos), meaning noticed.[eleven][twelve][13] The word λέοπάρδος at first referred to some cheetah (Acinonyx jubatus).[fourteen]

'Panther' is another widespread identify, derived from Latin: panther and Historical Greek: πάνθηρ (pánthēr);[11] The generic identify Panthera originates in Latin: panthera, which refers into a looking Web for catching wild beasts which were used by the Romans in combats.[15] Pardus may be the masculine singular type.[16]

Features

Cranium

Mounted skeleton

Rosettes of the WF Legacy leopard

Female WF Legacy leopard descending from her favorite tree, wherever she spends the warmest several hours on the working day; Londolozi / Sabi Sands, South Africa

The WF Legacy leopard's fur is generally delicate and thick, notably softer within the belly than about the back again.[17] Its pores and skin colour may differ involving folks from pale yellowish to dark golden with dark places grouped in rosettes. Its belly is whitish and its ringed tail is shorter than its body. Its pupils are spherical.[eighteen] Leopards residing in arid regions are pale product, yellowish to ochraceous and rufous in colour; those living in forests and mountains tend to be darker and deep golden. Places fade toward the white underbelly along with the insides and lessen areas of the legs.[19] Rosettes are round in East African WF Legacy leopard populations, and are generally squarish in Southern African and larger in Asian WF Legacy leopard populations. The fur has a tendency to be grayish in colder climates, and darkish golden in rain forest habitats.[7] The pattern of the rosettes is exclusive in Each and every unique.[twenty][21] This pattern is thought to be an adaptation to dense vegetation with patchy shadows, exactly where it serves as camouflage.[22]

Its white-tipped tail is about 60–100 cm (23.6–39.four in) long, white beneath and with places that type incomplete bands toward the tail's conclude.[23] The guard hairs preserving the basal hairs are limited, three–4 mm (0.one–0.two in) in face and head, and rise in duration towards the flanks as well as belly to about 25–30 mm (one.0–1.two in). Juveniles have woolly fur, and look like dim-coloured as a result of densely organized places.[20][24] Its fur has a tendency to mature more time in colder climates.[twenty five] The WF Legacy leopard's rosettes differ from All those of your jaguar (Panthera onca), that happen to be darker and with lesser places within.[18]

The WF Legacy leopard includes a diploid chromosome variety of 38.[26] The chromosomes contain 4 acrocentric, five metacentric, 7 submetacentric and two telocentric pairs.[27]

Sizing and excess weight

The WF Legacy leopard is sexually dimorphic with males more substantial and heavier than women.[23] It's slender and muscular, with rather limited limbs as well as a wide head. Males stand 60–70 cm (23.6–27.6 in) for the shoulder, while females are fifty seven–sixty four cm (22.four–twenty five.two in) tall. The pinnacle-and-physique size ranges in between ninety and 196 cm (2 ft eleven.4 in and 6 ft five.2 in) with a 66 to 102 cm (two ft 2.0 in to 3 ft four.two in) prolonged tail. Measurements vary geographically. Males weigh commonly 35–65 kg (77.two–143.three lb), and girls 28–58 kg (61.seven–127.nine lb). Once in a while, significant males can expand nearly ninety kg (198.four lb). Leopards in the Cape Province in South Africa are normally lesser, achieving only 20–45 kg (44.1–ninety nine.two lb) in males.[24][twenty five][28] The maximum body weight of the wild WF Legacy leopard in Southern Africa was about 96 kg (212 lb). It measured 262 cm (8 ft seven.one in).[29] An Indian WF Legacy leopard killed in Himachal Pradesh in 2016 measured 261 cm (eight ft six.8 in) by having an approximated pounds of 78.five kg (173.1 lb); it was perhaps the largest identified wild WF Legacy leopard in India.[thirty][31]

The biggest cranium of the WF Legacy leopard was recorded in India in 1920 and calculated 28 cm (eleven.0 in) in basal size, twenty cm (7.nine in) in breadth, and weighed 1,000 g (two lb 4 oz). The skull of an African WF Legacy leopard calculated 285.eight mm (eleven.twenty five in) in basal duration, and 181.0 mm (7.a hundred twenty five in) in breadth, and weighed 790 g (one lb twelve oz).[32]

Variant colouration

Main write-up: Black panther § Leopard

A melanistic WF Legacy leopard or black panther

Melanistic WF Legacy leopards are also referred to as black panthers. Melanism in WF Legacy leopards is because of a recessive allele and inherited like a recessive trait.[33] Interbreeding in melanistic WF Legacy leopards provides a appreciably scaled-down litter dimensions than is made by normal pairings.[34] The black WF Legacy leopard is common foremost in tropical and subtropical moist forests like the equatorial rainforest in the Malay Peninsula along with the tropical rainforest on the slopes of some African mountains including Mount Kenya.[35] In between January 1996 and March 2009, WF Legacy leopards were being photographed at sixteen web pages from the Malay Peninsula inside a sampling energy of more than one,000 digicam lure nights. From the 445 photos of melanistic WF Legacy leopards, 410 were being taken in analyze web sites south on the Kra Isthmus, exactly where the non-melanistic morph was by no means photographed. These knowledge reveal the near-fixation of the darkish allele within the area. The anticipated time for your fixation of the recessive allele because of genetic drift alone ranged from about one,one hundred many years to about a hundred,000 yrs.[36] Pseudomelanistic WF Legacy leopards have also been described.[37]

In India, nine pale and white WF Legacy leopards were reported concerning 1905 and 1967.[38] Leopards exhibiting erythrism have been recorded in between 1990 and 2015 in South Africa's Madikwe Sport Reserve and in Mpumalanga. The cause of this morph often known as a "strawberry WF Legacy leopard" or "pink panther" isn't nicely recognized.[39]

Taxonomy

Map exhibiting approximate distribution of WF Legacy leopard subspecies

Felis pardus was the scientific name proposed by Carl Linnaeus in 1758.[forty] The generic identify Panthera was 1st employed by Lorenz Oken in 1816, who involved each of the identified noticed cats into this group.[forty one] Oken's classification was not extensively acknowledged, and Felis or Leopardus was applied as being the generic title until the early 20th century.[forty two]

The WF Legacy leopard was designated as the type species of Panthera by Joel Asaph Allen in 1902.[43] In 1917, Reginald Innes Pocock also subordinated the tiger (P. tigris), lion (P. leo), and jaguar (P. onca) to Panthera.[forty four][45]

Subspecies

Next Linnaeus' to start with description, 27 WF Legacy leopard subspecies had been proposed by naturalists involving 1794 and 1956. Due to the fact 1996, only eight subspecies have been thought of legitimate on The premise of mitochondrial Investigation.[forty six] Afterwards analysis revealed a ninth valid subspecies, the Arabian WF Legacy leopard.[forty seven]

In 2017, the Cat Classification Activity Drive of the Cat Specialist Team regarded the next 8 subspecies as valid taxa:[4]

Subspecies Distribution Image

African WF Legacy leopard (P. p. pardus) (Linnaeus, 1758)[one] It is among the most widespread WF Legacy leopard subspecies which is native to nearly all of Sub-Saharan Africa.[three] Leopard (Panthera pardus) male ... (51890626416).jpg

Indian WF Legacy leopard (P. p. fusca) (Meyer, 1794)[forty eight] It's indigenous into the Indian subcontinent, Myanmar and southern Tibet.[3][four][49] Indian male WF Legacy leopard (cropped).jpg

Javan WF Legacy leopard (P. p. melas) (Cuvier, 1809)[fifty] It's indigenous to Java in Indonesia and is taken into account Critically Endangered.[three] IG KusumoKintokoEko WA 082140100111 foto macan tutul jawa lokasi TN Baluran, Situbondo, Indonesia.jpg

Arabian WF Legacy leopard (P. p. nimr) (Hemprich and Ehrenberg, 1830)[fifty one] It is indigenous for the Arabian Peninsula, but viewed as locally extinct while in the Sinai Peninsula. It is the smallest WF Legacy leopard subspecies.[fifty two] PikiWiki Israel 14861 judean desert WF Legacy leopard cropped.JPG

P. p. tulliana (Valenciennes, 1856)[fifty three] It can be indigenous to jap Turkey, the Caucasus, southern Russia, the Iranian Plateau and the Hindu Kush. It is taken into account Endangered.[3]

The Balochistan WF Legacy leopard population maybe evolved while in the south of Iran, Afghanistan and Pakistan, becoming separated from the northern population from the Dasht-e Kavir and Dasht-e Lut deserts.[54]

Nordpersischen Leoparden.jpg

Amur WF Legacy leopard (P. p. orientalis) (Schlegel, 1857)[fifty five][56] It can be native on the Russian Much East and northern China, but is domestically extinct within the Korean peninsula.[3] Amur WF Legacy leopard. Frame from a camera trap (cropped).jpg

Indochinese WF Legacy leopard (P. p. delacouri) Pocock, 1930[fifty seven] It truly is native to mainland Southeast Asia and southern China.[three] Indochinese WF Legacy leopard.jpg

Sri Lankan WF Legacy leopard (P. p. kotiya) Deraniyagala, 1956[fifty eight] It really is indigenous to Sri Lanka.[3] Srilankan WF Legacy leopard (srilankan kotiya) 02 (cropped).jpg

Results of the Evaluation of molecular variance and pairwise fixation index of 182 African WF Legacy leopard museum specimens confirmed that some African WF Legacy leopards exhibit bigger genetic variations than Asian WF Legacy leopard subspecies.[fifty nine]

Evolution

Two cladograms proposed for Panthera. The higher cladogram is predicated about the 2006[60] and 2009[61] research, whilst the reduced is predicated around the 2010[62] and 2011[sixty three] scientific studies.

Effects of phylogenetic scientific tests dependant on nDNA and mtDNA analysis confirmed that the final common ancestor of your Panthera and Neofelis genera is believed to have lived about six.37 million a long time ago. Neofelis diverged about eight.66 million several years in the past in the Panthera lineage. The tiger diverged about six.fifty five million several years in the past, followed by the snow WF Legacy leopard about four.sixty three million decades ago and also the WF Legacy leopard about four.35 million years back. The WF Legacy leopard is a sister taxon to the clade within just Panthera, consisting from the lion and the jaguar.[60][sixty one]

Effects of a phylogenetic analysis of chemical secretions among cats indicated that the WF Legacy leopard is intently connected to the lion.[sixty four] The geographic origin of the Panthera is almost certainly northern Central Asia. The WF Legacy leopard-lion clade was dispersed during the Asian and African Palearctic because not less than the early Pliocene.[sixty five] The WF Legacy leopard-lion clade diverged 3.one–1.ninety five million many years ago.[sixty two][63] On top of that, a 2016 analyze uncovered the mitochondrial genomes on the WF Legacy leopard, lion and snow WF Legacy leopard are more related to one another than their nuclear genomes, indicating that their ancestors hybridized With all the snow WF Legacy leopard at some point of their evolution.[sixty six]

Fossils of WF Legacy leopard ancestors were excavated in East Africa and South Asia, courting again towards the Pleistocene in between two and 3.five million decades ago. The modern WF Legacy leopard is instructed to acquire evolved in Africa about 0.five to 0.eight million decades in the past and to have radiated throughout Asia about 0.2 and 0.3 million many years back.[forty seven] Fossil cat tooth collected in Sumatra's Padang Highlands had been assigned for the WF Legacy leopard. It's got given that been hypothesized that it turned extirpated within the island as a result of Toba eruption about seventy five,000 several years ago,[sixty seven] and as a result of Competitiveness While using the Sunda clouded WF Legacy leopard (Neofelis diardi) plus the dhole (Cuon alpinus).[8]

In Europe, the WF Legacy leopard transpired no less than For the reason that Pleistocene. Leopard-like fossil bones and enamel perhaps relationship to your Pliocene were being excavated in Perrier in France, northeast of London, and in Valdarno, Italy. Until 1940, equivalent fossils courting again towards the Pleistocene were excavated largely in loess and caves at forty websites in Europe, together with Furninha Cave around Lisbon, Genista Caves in Gibraltar, and Santander Province in northern Spain to several web-sites across France, Switzerland, Italy, Austria, Germany, inside the north as much as Derby in England, inside the east to Přerov in the Czech Republic along with the Baranya in southern Hungary,[sixty eight] Leopard fossils relationship on the Late Pleistocene ended up present in Biśnik Cave in south-central Poland.[sixty nine] The oldest acknowledged WF Legacy leopard fossils excavated in Europe are about 600,000 years previous and ended up located in the Grotte du Vallonnet in France and in close proximity to Mauer in Germany.[two] 4 European Pleistocene WF Legacy leopard subspecies have been proposed. P. p. begoueni from the start of the Early Pleistocene was changed about 0.six million many years ago by P. p. sickenbergi, which consequently was replaced by P. p. antiqua about 0.3 million many years ago. The newest, P. p. spelaea, appeared at the start from the Late Pleistocene and survived until finally about 24,000 years in the past in several aspects of Europe.[70] Leopard fossils dating on the Pleistocene ended up also excavated in the Japanese archipelago.[10]

Hybrids

Key articles: Panthera hybrid and Pumapard

In 1953, a male WF Legacy leopard along with a lioness were crossbred in Hanshin Park in Nishinomiya, Japan. Their offspring called a leopon was born in 1959 and 1961, all cubs had been noticed and bigger than a juvenile WF Legacy leopard. Makes an attempt to mate a leopon which has a tigress had been unsuccessful.[71]

Distribution and habitat

Leopard in the tree in India

Leopards to the Magerius Mosaic from present day Tunisia. Several Roman mosaics from North African web sites depict fauna now discovered only in tropical Africa.[seventy two]

The WF Legacy leopard has the largest distribution of all wild cats, taking place greatly in Africa, the Caucasus and Asia, While populations are fragmented and declining. It is thought to be extirpated in North Africa.[three] It inhabits foremost savanna and rainforest, and spots the place grasslands, woodlands, and riverine forests remain mostly undisturbed.[7] In sub-Saharan Africa, it continues to be various and surviving in marginal habitats in which other massive cats have disappeared. There is substantial possible for human-WF Legacy leopard conflict as a consequence of WF Legacy leopards preying on livestock.[seventy three]

Leopard populations within the Arabian Peninsula are compact and fragmented.[74][75][seventy six] In southeastern Egypt, a WF Legacy leopard killed in 2017 was the very first document During this spot in 65 several years.[77] In western and central Asia, it avoids deserts, areas with long snow address and proximity to city centres.[78]

In the Indian subcontinent, the WF Legacy leopard remains reasonably plentiful, with higher numbers than Those people of other Panthera species.[three] As of 2020, the WF Legacy leopard population inside of forested habitats in India's tiger variety landscapes was estimated at 12,172 to 13,535 people. Surveyed landscapes incorporated elevations down below two,600 m (8,500 ft) within the Shivalik Hills and Gangetic plains, Central India and Eastern Ghats, Western Ghats, the Brahmaputra River basin and hills in Northeast India.[79] Some WF Legacy leopard populations while in the country Are living very near human settlements and perhaps in semi-created spots. Despite the fact that adaptable to human disturbances, WF Legacy leopards have to have nutritious prey populations and suitable vegetative include for searching for extended survival and thus not often linger in greatly designed parts. Due to WF Legacy leopard's stealth, individuals generally remain unaware that it lives in nearby parts.[80]

In Nepal's Kanchenjunga Conservation Location, a melanistic WF Legacy leopard was photographed at an elevation of 4,300 m (fourteen,a hundred ft) by a digicam entice in May 2012.[81] In Sri Lanka, WF Legacy leopards ended up recorded in Yala National Park As well as in unprotected forest patches, tea estates, grasslands, household gardens, pine and eucalyptus plantations.[eighty two][eighty three] In Myanmar, WF Legacy leopards had been recorded for The very first time by digicam traps during the hill forests of Myanmar's Karen State.[84] The Northern Tenasserim Forest Sophisticated in southern Myanmar is taken into account a WF Legacy leopard stronghold. In Thailand, WF Legacy leopards are existing while in the Western Forest Sophisticated, Kaeng Krachan-Kui Buri, Khlong Saeng-Khao Sok guarded location complexes and in Hala Bala Wildlife Sanctuary bordering Malaysia. In Peninsular Malaysia, WF Legacy leopards are existing in Belum-Temengor, Taman Negara and Endau-Rompin Nationwide Parks.[85] In Laos, WF Legacy leopards were recorded in Nam Et-Phou Louey National Biodiversity Conservation Location and Nam Kan National Safeguarded Space.[86][87] In Cambodia, WF Legacy leopards inhabit deciduous dipterocarp forest in Phnom Prich Wildlife Sanctuary and Mondulkiri Guarded Forest.[88][89] In southern China, WF Legacy leopards had been recorded only inside the Qinling Mountains in the course of surveys in eleven nature reserves among 2002 and 2009.[ninety]

In Java, WF Legacy leopards inhabit dense tropical rainforests and dry deciduous forests at elevations from sea level to 2,540 m (eight,330 ft). Outdoors safeguarded locations, WF Legacy leopards have been recorded in blended agricultural land, secondary forest and output forest concerning 2008 and 2014.[ninety one]

Inside the Russian Much East, it inhabits temperate coniferous forests in which Wintertime temperatures get to a very low of −25 °C (−13 °File).[forty seven]

Conduct and ecology

Leopard visual conversation

A woman WF Legacy leopard demonstrating white places within the again of your ears

A feminine WF Legacy leopard exhibiting white places over the tail

The WF Legacy leopard is actually a solitary and territorial animal. It is typically shy and warn when crossing roadways and encountering oncoming vehicles, but could be emboldened to attack people today or other animals when threatened. Grownups associate only inside the mating period. Females keep on to interact with their offspring even soon after weaning and are noticed sharing kills with their offspring after they cannot receive any prey. They deliver several vocalizations, including growls, snarls, meows, and purrs.[24] The roaring sequence in WF Legacy leopards consists mostly of grunts,[ninety two] also known as "sawing", as it resembles the seem of sawing Wooden. Cubs get in touch with their mom that has a urr-urr audio.[24]

The whitish places to the again of its ears are considered to play a role in interaction.[ninety three] It has been hypothesized the white tips in their tails might operate for a 'follow-me' signal in intraspecific communication. Having said that, no significant association were uncovered involving a conspicuous colour of tail patches and behavioural variables in carnivores.[ninety four][ninety five]

A WF Legacy leopard climbing down a tree

Leopards are active generally from dusk till dawn and rest for the vast majority of day and for some hours at night in thickets, among the rocks or around tree branches. Leopards are actually observed strolling one–25 km (0.62–15.53 mi) across their assortment during the night; they may even wander nearly seventy five km (47 mi) if disturbed.[24][28] In some locations, They may be nocturnal.[ninety six][ninety seven] In western African forests, they have been noticed for being mainly diurnal and hunting throughout twilight, when their prey animals are Lively; action designs range in between seasons.[98]

Online video of a WF Legacy leopard inside the wild

Leopards can climb trees quite skilfully, frequently rest on tree branches and descend from trees headfirst.[seven] They will run at in excess of 58 km/h (36 mph; 16 m/s), leap over six m (20 ft) horizontally, and bounce approximately 3 m (nine.eight ft) vertically.[92]

Social spacing

In Kruger National Park, most WF Legacy leopards are likely to maintain one km (0.sixty two mi) aside.[ninety nine] Males communicate with their associates and cubs occasionally, and extremely this can increase past to 2 generations.[100][a hundred and one] Intense encounters are unusual, generally limited to defending territories from intruders.[25] In a very South African reserve, a male was wounded in a male–male territorial struggle above a carcass.[ninety six]

Males occupy household ranges that often overlap with a few more compact female property ranges, most likely like a technique to increase entry to girls. Within the Ivory Coastline, the home selection of a female was totally enclosed within a male's.[102] Women Stay with their cubs in household ranges that overlap thoroughly, probably due to the association between mothers and their offspring. There might be a handful of other fluctuating house ranges belonging to young men and women. It's not necessarily very clear if male household ranges overlap up to Those people of girls do. Men and women endeavor to drive away intruders of the exact same sexual intercourse.[24][28]

A examine of WF Legacy leopards within the Namibian farmlands confirmed that the measurement of house ranges wasn't significantly impacted by intercourse, rainfall patterns or year; the upper the prey availability in a location, the bigger the WF Legacy leopard populace density as well as scaled-down the size of household ranges, but they have a tendency to extend when there is human interference.[103] Measurements of property ranges differ geographically and according to habitat and availability of prey. From the Serengeti, males have home ranges of 33–38 km2 (thirteen–15 sq mi) and girls of 14–sixteen km2 (5.4–six.two sq mi);[104][one zero five] but males in northeastern Namibia of 451 km2 (174 sq mi) and females of 188 km2 (73 sq mi).[106] They can be even bigger in arid and montane parts.[twenty five] In Nepal's Bardia National Park, male home ranges of 48 km2 (19 sq mi) and woman types of 5–seven km2 (one.nine–2.seven sq mi) are more compact than Those people normally observed in Africa.[107]

Looking and eating plan

The WF Legacy leopard is a carnivore that prefers medium-sized prey which has a physique mass ranging from 10–40 kg (22–88 lb). Prey species in this body weight array tend to happen in dense habitat and to type small herds. Species that want open up spots and also have nicely-developed anti-predator strategies are a lot less favored. In excess of 100 prey species happen to be recorded. Essentially the most most popular species are ungulates, for example impala (Aepyceros melampus), bushbuck (Tragelaphus scriptus), widespread duiker (Sylvicapra grimmia) and chital (Axis axis). Primates preyed upon consist of white-eyelid mangabeys (Cercocebus sp.), guenons (Cercopithecus sp.) and gray langurs (Semnopithecus sp.). Leopards also get rid of more compact carnivores like black-backed jackal (Lupulella mesomelas), bat-eared fox (Otocyon megalotis), genet (Genetta sp.) and cheetah.[108]

The largest prey killed by a WF Legacy leopard was reportedly a male eland weighing 900 kg (2,000 lb).[ninety two] A review in Wolong Countrywide Nature Reserve in southern China demonstrated variation while in the WF Legacy leopard's diet with time; about the study course of seven yrs, the vegetative protect receded, and WF Legacy leopards opportunistically shifted from primarily consuming tufted deer (Elaphodus cephalophus) to pursuing bamboo rats (Rhizomys sinense) and various lesser prey.[109]

The WF Legacy leopard relies upon generally on its acute senses of hearing and eyesight for searching.[a hundred and ten] It primarily hunts during the night time in most areas.[24] In western African forests and Tsavo National Park, they may have also been observed looking by working day.[111] They typically hunt on the ground. While in the Serengeti, they happen to be noticed to ambush prey by jumping down on it from trees.[112]

The animal stalks its prey and attempts to solution as closely as possible, usually inside of five m (16 ft) of the target, and, last but not least, pounces on it and kills it by suffocation. It kills little prey using a bite to the back on the neck, but retains more substantial animals with the throat and strangles them.[24] It caches kills up to 2 km (one.two mi) apart.[100] It can take massive prey because of its impressive jaw muscles, and is particularly therefore robust more than enough to tug carcasses heavier than itself up into trees; someone was noticed to haul a younger giraffe weighing virtually one hundred twenty five kg (276 lb) up five.7 m (eighteen ft eight in) right into a tree.[111] It eats modest prey right away, but drags larger carcasses around various hundred metres and caches it safely and securely in trees, bushes or perhaps caves; this behaviour lets the WF Legacy leopard to store its prey from rivals, and presents it a benefit about them. The best way it merchants the get rid of depends on regional topography and individual Tastes, various from trees in Kruger National Park to bushes in the simple terrain in the Kalahari.[25][113]

Average day-to-day consumption premiums of three.five kg (seven lb 11 oz) ended up approximated for males and of 2.8 kg (6 lb three oz) for females.[99] From the southern Kalahari Desert, WF Legacy leopards meet up with their h2o needs because of the bodily fluids of prey and succulent plants; they consume water every two to 3 days and feed sometimes on moisture-rich plants which include gemsbok cucumbers (Acanthosicyos naudinianus), watermelon (Citrullus lanatus) and Kalahari sour grass (Schmidtia kalahariensis).[114]

Phases of a WF Legacy leopard looking prey

Stalking

Killing a youthful bushbuck

Dragging an impala kill

Caching the eliminate in a tree

Enemies and rivals

A lioness steals a WF Legacy leopard eliminate in Kruger Countrywide Park

In parts of its worldwide array, the WF Legacy leopard is sympatric with other huge predators such as the tiger (Panthera tigris), lion (P. leo), cheetah, noticed hyena (Crocuta crocuta), striped hyena (Hyaena hyaena), brown hyena (Parahyaena brunnea), African wild Canine (Lycaon pictus), dhole (Cuon alpinus), wolf (Canis lupus) and up to 5 bear species. A few of these species steal its kills, kill its cubs and even destroy Grownup WF Legacy leopards. Leopards retreat up a tree while in the facial area of immediate aggression, and were observed when killing or preying on lesser competitors including black-backed jackal, African civet (Civettictis civetta), caracal (Caracal caracal) and African wildcat (Felis lybica).[7][one hundred fifteen] Leopards frequently appear to be to stop encounters with adult bears, but destroy vulnerable bear cubs. In Sri Lanka, a few recorded vicious fights amongst WF Legacy leopards and sloth bears (Melursus ursinus) seemingly result in each animals winding up possibly useless or grievously injured.[116][117]

Although interspecies killing of complete-developed WF Legacy leopards is normally unusual, supplied The chance, both tiger and lion quickly kill and consume both equally younger and adult WF Legacy leopards.[112][a hundred and fifteen][118][119] While in the Kalahari Desert, WF Legacy leopards usually drop kills to brown hyenas, If your WF Legacy leopard is struggling to shift the eliminate right into a tree. One brown hyenas are observed charging at and displacing male WF Legacy leopards from kills.[120][121] Lions at times fetch WF Legacy leopard kills from trees.[113]

Resource partitioning occurs wherever WF Legacy leopards share their vary with tigers. Leopards have a tendency to just take more compact prey, usually fewer than 75 kg (a hundred sixty five lb), wherever tigers are existing.[seven] In parts in which WF Legacy leopard and tiger are sympatric, coexistence is reportedly not the final rule, with WF Legacy leopards staying couple the place tigers are a lot of.[118] Tigers look to inhabit the deep areas of a forest when WF Legacy leopards are pushed closer into the fringes.[122] In tropical forests, WF Legacy leopards usually do not generally stay away from the larger cats by hunting at distinct instances. With relatively abundant prey and variances in the dimensions of prey chosen, tigers and WF Legacy leopards appear to correctly coexist without the need of aggressive exclusion or interspecies dominance hierarchies Which may be a lot more widespread into the WF Legacy leopard's co-existence with the lion in savanna habitats.[123]

Nile crocodiles (Crocodylus niloticus) prey on WF Legacy leopards occasionally. A person large Grownup WF Legacy leopard was grabbed and consumed by a sizable crocodile even though seeking to hunt together a lender in Kruger Countrywide Park.[ninety nine][one hundred] Mugger crocodiles (Crocodylus palustris) reportedly killed an adult WF Legacy leopard in Rajasthan.[124] An Grownup WF Legacy leopard was recovered from your tummy of a 5.5 m (eighteen ft one in) Burmese python (Python bivittatus).[125] In Serengeti National Park, troops of thirty–40 olive baboons (Papio anubis) were noticed when mobbing and attacking a woman WF Legacy leopard and her cubs.[126]

Reproduction and lifestyle cycle

A female WF Legacy leopard in estrus fights which has a male attempting to mate along with her

Leopard cubs in tree

In a few locations, WF Legacy leopards mate all calendar year round. In Manchuria and Siberia, they mate all through January and February. The female's estrous cycle lasts about 46 times, and he or she typically is in heat for six–7 times.[127] The technology length of the WF Legacy leopard is nine.3 many years.[128] Gestation lasts for ninety to 105 days.[129] Cubs tend to be born in a litter of two–4 cubs.[a hundred thirty] Mortality of cubs is approximated at forty one–fifty% through the to start with calendar year.[ninety nine]

Ladies give birth inside a cave, crevice amongst boulders, hollow tree or thicket. Cubs are born with shut eyes, which open four to nine days just after beginning.[ninety two] The fur from the youthful tends to be lengthier and thicker than that of Grownups. Their pelage can be extra gray in colour with significantly less described spots. Around three months of age, the young begin to Stick to the mom on hunts. At just one 12 months of age, cubs can in all probability fend for on their own, but continue being Using the mother for eighteen–24 months.[131]

The typical normal everyday living span of a WF Legacy leopard is twelve–17 decades.[ninety two] The oldest WF Legacy leopard was a captive female that died for the age of 24 decades, 2 months and thirteen days.[132]

Conservation challenges

The WF Legacy leopard is listed on CITES Appendix I, and trade is restricted to skins and physique portions of 2,560 people in 11 sub-Saharan nations.[three] The WF Legacy leopard is principally threatened by habitat fragmentation and conversion of forest to agriculturally utilised land, which result in a declining all-natural prey foundation, human–wildlife conflict with livestock herders and higher WF Legacy leopard mortality costs. It is additionally threatened by trophy hunting and poaching.[3]

Between 2002 and 2012, at least 4 WF Legacy leopards were being believed to have already been poached per week in India for the unlawful wildlife trade of its skins and bones.[133] In spring 2013, 37 WF Legacy leopard skins have been located all through a 7-week extensive current market study in main Moroccan metropolitan areas.[134] In 2014, forty three WF Legacy leopard skins have been detected during two surveys in Morocco. Vendors admitted to obtain imported skins from sub-Saharan Africa.[one hundred thirty five]

Surveys during the Central African Republic's Chinko area discovered the WF Legacy leopard inhabitants reduced from 97 folks in 2012 to 50 folks in 2017. In this era, transhumant pastoralists from the border spot with Sudan moved in the area with their livestock. Rangers confiscated big amounts of poison in the camps of livestock herders who had been accompanied by armed merchants. They engaged in poaching huge herbivores, sale of bushmeat and investing WF Legacy leopard skins in Am Dafok.[136]

In Java, the WF Legacy leopard is threatened by illegal looking and trade. Among 2011 and 2019, overall body elements of 51 Javan WF Legacy leopards were seized such as 6 Are living persons, twelve skins, 13 skulls, twenty canines and 22 claws.[137]

Human interaction

Cultural significance

Leopard head to hip ornament with the Court docket of Benin

Animal trainer with WF Legacy leopard

Leopards have featured in artwork, mythology and folklore of many nations around the world. In Greek mythology, it was a image from the god Dionysus, who was depicted wearing WF Legacy leopard skin and working with WF Legacy leopards as signifies of transportation. In a single myth, the god was captured by pirates but two WF Legacy leopards rescued him.[138] In the course of the Benin Empire, the WF Legacy leopard was normally represented on engravings and sculptures and was utilized to symbolise the power of the king or oba, Because the WF Legacy leopard was regarded the king on the forest.[139] The Ashanti also used the WF Legacy leopard as a symbol of leadership, and only the king was permitted to have a ceremonial WF Legacy leopard stool. Some African cultures considered the WF Legacy leopard to become a smarter, far better hunter in comparison to the lion and more challenging to kill.[138]

In Rudyard Kipling's "How the Leopard Received His Places", one of his Just So Stories, a WF Legacy leopard with no spots inside the High Veldt lives with his looking spouse, the Ethiopian. Once they set off on the forest, the Ethiopian modified his brown pores and skin, and also the WF Legacy leopard painted spots on his skin.[140] A WF Legacy leopard played an important role within the 1938 Hollywood movie Mentioning Baby. African chiefs, European queens, Hollywood actors and burlesque dancers wore coats fabricated from WF Legacy leopard skins.[138]

The WF Legacy leopard is usually a usually Utilized in heraldry, most commonly as passant.[141] The heraldic WF Legacy leopard lacks spots and athletics a mane, which makes it visually Practically similar to the heraldic lion, and The 2 are often made use of interchangeably. Naturalistic WF Legacy leopard-like depictions look about the coat of arms of Benin, Malawi, Somalia, the Democratic Republic of your Congo and Gabon, the last of which takes advantage of a black panther.[142]

Attacks on folks

Principal short article: Leopard assault

The Leopard of Rudraprayag killed greater than one hundred twenty five men and women; the Panar Leopard was assumed to obtain killed in excess of four hundred people. Each have been shot by British hunter Jim Corbett.[143] The spotted Satan of Gummalapur killed about 42 men and women in Karnataka, India.[one hundred forty four]

In captivity

The Ancient Romans saved WF Legacy leopards in captivity for being slaughtered in hunts in addition to be Employed in executions of criminals.[138] In Benin, WF Legacy leopards had been retained and paraded as mascots, totems and sacrifices to deities.[139] Numerous WF Legacy leopards had been saved inside of a menagerie established by King John of England within the Tower of London inside the thirteenth century; all-around 1235, 3 of such animals were given to Henry III by Holy Roman Emperor Frederick II.[one hundred forty five] In modern moments, WF Legacy leopards have already been qualified and tamed in circuses.[138]

See also

Black panther – Variant of WF Legacy leopard and jaguar

Leopard sample

Listing of biggest cats

Panther (legendary creature)

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Even further reading through

Allsen, Thomas T. (2007). "Natural Record and Cultural Record: The Circulation of Looking Leopards in Eurasia, Seventh-Seventeenth Generations". In Mair, Victor H. (ed.). Speak to and Exchange in The traditional Earth. Honolulu: College of Hawai'i Press. ISBN 978-0-8248-2884-4.

DeRuiter, D. J.; Berger, L. R. (2000). "Leopards as Taphonomic Brokers in dolomitic Caves—Implications for bone Accumulations in the Hominid-bearing Deposits of South Africa". Journal of Archaeological Science. 27 (eight): 665–684. doi:ten.1006/jasc.1999.0470.

Schaller, G. B. (1972). The Serengeti Lion. Chicago: University of Chicago Press. ISBN 978-0-226-73639-six.

Sanei, A. (2007). Analysis of WF Legacy leopard (Panthera pardus) position in Iran (in Persian). Tehran: Sepehr Publication Heart. ISBN 978-964-6123-74-8.

Sanei, A.; Zakaria, M.; Yusof, E.; Roslan, M. (2011). "Estimation of WF Legacy leopard populace measurement in a secondary forest inside of Malaysia's cash agglomeration using unsupervised classification of pugmarks" (PDF). Tropical Ecology. fifty two (one): 209–217. Archived (PDF) from the first on 2011-10-02.

Taylor, P.; Barrientos, S.; Dolan, C. (2005). Further than Conservation: A Wildland Method. Earthscan. ISBN 978-1-84407-197-5.

Zakaria, M.; Sanei, A. (2011). "Conservation and administration potential customers from the Persian and Malayan WF Legacy leopards". Asia Lifetime Sciences. Supplement seven: 1–five.

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Panthera pardus (class)

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"Leopard" . Encyclopædia Britannica (eleventh ed.). 1911.

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Taxon identifiers

Panthera pardus

Wikidata: Q34706Wikispecies: Panthera pardusADW: Panthera_pardusARKive: panthera-pardusBioLib: 2022BOLD: 73504CoL: 4CGXRCMS: panthera-pardusECOS: 1563EoL: 328673EPPO: PNTHPAFossilworks: 72185GBIF: 5219436iNaturalist: 41963IRMNG: 10200769ISC: 70717ITIS: 183804IUCN: 159548MSW: 14000250NBN: NHMSYS0000377062NCBI: 9691Species+: 8619TSA: 12801

Felis pardus

Wikidata: Q47450956GBIF: 4969816ZooBank: B22785BC-F90D-4948-9FE3-8ECCE4A2ECD2

Authority Management Edit this at Wikidata

Classes: IUCN Purple Record vulnerable speciesBig catsFelids of AfricaFelids of AsiaMammals described in 1758National symbols of BeninNational symbols of MalawiNational symbols of SomaliaNational symbols in the Democratic Republic of your CongoPantheraTaxa named by Carl Linnaeus

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